There are also two other genetic "issues" to consider... the first is spontaneous mutation - for additional information on this Google "Panda SHepherds" - pure West German highline GSDs that produced white-patched pups - one parent had a spontaneous mutation for this and it is currently being studied by University of California. ANother is "Chimerism" - Google "Chimera" where unexpected DNA plays a role - brindle colored horses, for instance, and it has been seen in humans, where a woman gave birth to her SISTER'S offspring (the baby she concieved and gave birth to was NOT genetically related to her but to a "sister" whose genes she carries as the "sister" was absorbed by her in the fetal stage).
Just because wild wolves, GSDs, etc. do not display mutations of color does not mean they do not exist in the gene pool.
Remember all dogs descend from the wolf, so there is the potential for anything in the genetics. Pushed-in faces are not seen in the wolf or GSD but is seen in wolf descendents like the Bully breeds, showing that wolf genetics are indeed a maleable "material" and likely to have spontaneous mutations from time to time...
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